Emma Swan, a 28-year-old bail bonds collector, has always been a fiercely independent person since being abandoned as a baby. Her son Henry, who she gave up for adoption years ago, finds and tries to convince Emma that she is Snow White's missing daughter. Henry shows Emma that in the fairytale, Prince Charming and Snow White sent her away to protect her. Emma doesn't believe him and takes Henry back to Storybrooke...

The Previously unknown purpose of the "Ninth Chevron" is revealed, and ends up taking a team to an Ancient ship "Destiny", a ship built millions of years ago by the Ancients, used to seed Distant galaxies with Stargates. This team, led by Dr. Nicolas Rush and Colonel Everett Young, is trapped on the ship, unable to change its programmed mission. The team encounters new races, new technology and new enemies as the runaway ship travels to the far ends of the Universe.